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Is the titan hand(starting at 0:02) animated on ones? It's very impressive.

Does anyone know for cuts animated on 1s, does the key animator usually do all the drawings or does a 2nd KA or inbetweener(not sure on the terminology) fill in the gaps?

It probably depends but I imagine it's the latter since having the main key animator do all that is a bit of a waste of resources unless they're very protective of their work.